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It is speculated that meiosis occurs during the time period from these formation of the conchospore to the first cell division of it. 竝對減數分裂這一遺傳現象進行了重新認識，提出紫菜減數分裂可能發生在殼孢子形成至殼孢子萌發的第一次細胞分裂時期。